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The Assassins concluded that the Templar-influenced ruler of the Roman Empire, Gaius Julius Caesar Augustus Germanicus, commonly referred to as Caligula, was a tyrant and the land had to be freed of him. On 24 January, 41 A.D., Leonius stabbed Caligula with a dagger in an underground corridor beneath Palatine Hill, killing him.
Between the time of his death and the Renaissance, Leonius' sarcophagus had been moved to Venice, Italy, beneath Santa Maria della Visitazione. The Assassin Ezio Auditore da Firenze explored Leonius' tomb and took the hidden seal from his sarcophagus, which would help him to later obtain Altaïr Ibn-La'Ahad's armor.