Jonathan Hawk is a member of the Assassin Order and a descendant of the Aleman Assassin Accipiter and Egyptian Assassin Numa Al'Khamsin. In 2012, he went into the Animus to relive the memories of his ancestors.
While in the Animus, he lived through the memories of his ancestor Accipiter of the Battle of Mediolanum in 259, between the Alemanni and the Romans in Aube. After spending a while longer with Accipiter's memories, Jonathan's nurse unplugged him from the Animus to let him rest for two hours, as a longer session could be dangerous for his mental health.
Later on, he visited Desmond Miles and the other Assassins in Monteriggioni to ask them for the "Folder Nº24" that Stella Crow had given to Desmond. In the process, he discovered he was a distant cousin to Desmond and, upon looking at the folder, Jonathan noticed a picture of an Egyptian pyramid, quickly realizing where his mission would take him next.